Samples
Sample 1. Khoe-San
Although sub-Saharan African groups are classified as being warm-adapted, low latitude populations, archaeological evidence suggests that the eco-geographic area inhabited by Khoe-Sans (descendants of the LSA hunter-gatherers who occupied the area) is significantly different. While the South African Cape is well below the North and South latitude division, the coastal climate is Mediterranean-like rather than tropical. The sample of 124 LSA hunter-gatherer skeletons (59 male and 65 female) from the Southern, Western, and Eastern Cape regions are dated between 240 and 7, 853 years BP.
From the tropical monsoon climate of the Andaman Island, 31 proto-historic Andaman Island forager skeletons (16 male and 15 female) were collected. These skeletons date to around the mid 1800s and originated from the Greater Andaman Island which is situated between latitude 11° and 14° north.
Sample 3.
Kurki et al. use published literature to collect data representing skeleton samples from Northern and Southern Europe, North Africa, and Eastern and Western Sub-Saharan Africa.
Kurki et al. tested 3 hypotheses to explain the relationship between latitude, climate, and biological adaptations: (1) if the body proportions observed among LSA hunter-gatherers reflect those of populations inhabiting low latitude and warm climate regions (e.g. high limb to skeleton trunk ratio), LSA hunter-gatherers of the South African Cape inhabited similar environments; (2) if the body proportions of the LSA hunter-gatherers are similar to those of the European and North African samples, the LSA hunter-gatherers likely inhabited a Mediterranean-like climate; and (3) if the body proportions resemble those of the Andaman foragers and African Pygmies, LSA hunter-gatherers were adapted to warm climates. Although Andaman foragers and African Pygmies are relatively small-bodied populations, their bi-iliac breadth to stature rations are equal to those of non small-bodied warm climate inhabiting populations.

Materials and Methods
Kurki et al. calculated the brachial, crural, femoral, humeral, radial, and tibial indices and body breadth and mass to stature using the ratios in table 1.
Limp Proportions:
Compared to the indices calculated for the 3 skeleton samples, the majority of the limb proportions of the LSA hunter-gatherers fall in between those of the North African and European samples.
Body Shape:
Proxy skeletal indicators were used to calculate stature ratios and the bi-iliac breadth determined the absolute body breadth. Kurki et al. found that body breadth decreased in relation to latitude, although the female African Pygmies have the largest bi-iliac breadth among the low latitude samples. Body mass also follows this trend, decreasing at lower latitudes. However, the Andaman Islander sample in which statures are similar to those of the African Pygmies have much more narrow bi-iliac breadths then expected. Sexual dimorphism was also observed in the LSA hunter-gatherer sample, although it was females who were generally larger then the males.
The results obtained in the study correspond to Kurki et al.’s third hypothesis. The skeletal morphology of the LSA hunter-gatherer sample falls between those of the European and North African Samples. Therefore, it appears that the Mediterranean-like climate of the southern African Cape had a greater influence on the LSA hunter-gatherer skeletal morphology than did the low latitude of the region. When compared to other African Sub-Saharan skeleton samples, the LSA hunter-gatherers and their descendants, the Khoe-San, have distinctly smaller limb proportions. From their findings, Kurki et al. caution against grouping populations into climate categories based on latitude. The eco-geographic patterning predictions on which Kurki et al. based this research was contradicted by the inconsistent body proportions of LSA hunter-gatherers and Khoe-San from the southern African Cape and the Andaman foragers.
Kurki et al.’s study of eco-geographic theories emphasizes that researchers consider the climate of regions inhabited by prehistoric populations. This is especially important when considering South African populations because the majority of archaeological sites are located on the coast. As demonstrated by Kurki et al., climates vary among and within latitudinal gradients. As a consequence, environmental adaptations may have developed for different climates then those predicted by latitude. However, the conclusions drawn from the study by Kurki et al. are based on methods that admittedly produce inaccurate results such as: “[…] the relationship between body mass and femoral height is not isometric, and the effect would be amplified in small bodied populations, such as those in this study” (p.30); There were several skeletons that were missing vertebral elements necessary to calculate skeleton trunk height. The height of these vertebral elements was estimated through a regression formula generated from a mixed-population reference sample which is rather problematic since the study sought to differentiate body proportions between populations; and only summary statistics were available for the skeleton samples obtained through published literature. Therefore, the samples were not directly comparable and a t-test identified differences in mean index value. The comparison of body proportions may have been somewhat altered. Finally, the indices and proxies for body proportions among different climates may be influenced by additional elements (e.g. life history parameters). Kurki et al. largely ignore the implications these additional elements may have on the body proportions of the sample populations in question.
Reference:
Kurki, H. K., J. K. Ginter, J. T. Stock, and S. Pfeiffer
2008 Adult Proportionality in Small-Bodied Foragers: A Test of Ecogeographic Expectations. American Journal of Physical Anthropology 136:28-38.